The World Flora Online

Arbres ou arbustes. Feuilles simples, alternes, entières ou dentées, stipulées (stipules caduques de petite taille). Inflorescences en grappes caulinaires ou en fascicules axillaires ou supra-axillaires. Fleurs ♂ ou ♀, actinomorphes, pentamères. Sépales imbriqués; pétales valvaires ou contortés. Étamines 5-10; anthères à deux loges introrses ou latérales, déhiscence longitudinale. Disque intrastaminal présent chez Panda ou absent. Ovaire rudimentaire dans la fleur♂, supère à 2-4 loges uniovulées dans la fleur ♀. Drupe à endocarpe lignifié, lacuneux et alvéolaire ou muriqué, rugueux. Graines 2-4 (-5).

Arbres'dioïques.'Feuilles'alternes, simples, à stipules petites et caduques.'Inflorescences'en racèmes caulinaires.'Fleurs'♂ fasciculées; calice cupulaire, petit et à préfloraison ouverte; corolle à 5 pétales valvaires; étamines 10 en 2 verticilles didynames, l'intérieur étant plus court que l'extérieur; disque intrastaminal peu développé; ovaire rudimentaire.'Fleurs'♀ solitaires; calice comme dans les fleurs ♂; corolle à 5 pétales ± imbriqués; staminodes nuls; gynécée à ovaire supère, subsessile, 3-4-loculaire, à 1 ovule orthotrope et pendant par loge; stigmates 2-4, subsessiles.'Fruits'drupacés, à mésocarpe bien développé; endocarpe épais, très dur, fortement alvéolé et lacuneux.'Graines 3-4, aplaties, à albumen oléagineux; embryon à larges cotylédons.\n\t\t\tUn seul genre monospécifique dans les forêts ombrophiles de l'Afrique tropicale occidentale.

Female flowers arranged like the male Stamens 10, alternately long and short; anthers 2-celled, opening lengthwise Rudimentary ovary linear-subulate Calyx cupular, open in bud Petals 5, valvate Flowers dioecious Male flowers racemose, the racemes fasciculate from the old wood Trees Leaves alternate, simple, serrulate; stipules present Ovary subsessile, 3–4-celled; stigmas 3–4, reflexed; ovule solitary, pendulous Fruit drupaceous Seeds with copious oily endosperm; cotyledons cordate Calyx cupular, truncate or toothed Petals 5 [imbricate?]; staminodes absent Stamens 10, alternately long and short; anthers 2-celled, opening lengthwise Rudimentary ovary linear-subulate Calyx cupular, open in bud Petals 5, valvate Flowers dioecious Male flowers racemose, the racemes fasciculate from the old wood Trees Leaves alternate, simple, serrulate; stipules present Ovary subsessile, 3–4-celled; stigmas 3–4, reflexed; ovule solitary, pendulous Fruit drupaceous Seeds with copious oily endosperm; cotyledons cordate Calyx cupular, truncate or toothed Petals 5 [imbricate?]; staminodes absent

Trees or shrubs, dioecious. Axillary buds prominent. Stipules small. Leaves alternate, distichous, petiolate, simple, penninerved, margin serrulate or entire. Flowers unisexual, solitary or fascicled, or in axillary cymes, or terminal or cauliflorous racemelike thyrses. Sepals 5, imbricate, or open. Petals 5, imbricate or valvate. Disk small or absent, rarely larger. Stamens 5-15, 1- or 2-seriate, sometimes unequal, outer stamens alternating with petals, inner ones sometimes absent or reduced to staminodes; filaments free; anthers introrse, thecae longitudinally dehiscent, connective elevated. Ovary superior, 2-5-loculed; ovules 1 or 2 per locule, apical, pendent, orthotropal or anatropal, without obturator; styles 2-5(-10)-lobed. Fruit a drupe, sometimes flattened, more rarely a capsule; endocarp slightly bony or pitted or ridged, sometimes dehiscent by valves. Seeds usually flattened concave, more rarely ovoid, caruncle absent; endosperm present; cotyledons broad and flattened. 2n = 30.

Dioecious trees or shrubs Leaves alternate, distichous, simple, entire, crenate or serrate, penninerved, stipulate, borne horizontally on shoots which subtend ‘axillary buds’ and thus resemble pinnate leaves; stipules usually persistent, the adaxial one arising below the abaxial Cotyledons broad, sometimes cordate Disc small, rarely large, or absent Petals 5, contorted, imbricate or valvate, flattened or galeate-cucullate Calyx cupular, truncate or 5-lobed, the lobes imbricate or open Flowers small, regular, unisexual, in axillary or supraaxillary fascicles, abbreviated racemes or cymes, or in terminal or cauliflorous racemiform thyrses Seeds usually flattened-concave, more rarely ovoid, ecarunculate, with endosperm Fruit a drupe, sometimes flattened, more rarely a capsule; mesocarp fleshy; endocarp woody, bony or stony, thin to thick and massive, usually tubercled, muricate, pitted or ridged, sometimes dehiscent by valves Ovary superior, 2–5-locular, with 1 (–2) apical pendulous orthotropous or anatropous ovules per locule, without obturator; styles 1–5, simple or bipartite Stamens 5, 10 or 15, uniseriate or biseriate, sometimes unequal; anthers introrse, 2-thecous, opening lengthwise; staminodes sometimes present in ? flowers; pistillode of ? flowers linear-subulate, cylindric or narrowly conical, sometimes pentagonal at base Leaves alternate, distichous, simple, entire, crenate or serrate, penninerved, stipulate, borne horizontally on shoots which subtend ‘axillary buds’ and thus resemble pinnate leaves; stipules usually persistent, the adaxial one arising below the abaxial Cotyledons broad, sometimes cordate Disc small, rarely large, or absent Petals 5, contorted, imbricate or valvate, flattened or galeate-cucullate Calyx cupular, truncate or 5-lobed, the lobes imbricate or open Flowers small, regular, unisexual, in axillary or supraaxillary fascicles, abbreviated racemes or cymes, or in terminal or cauliflorous racemiform thyrses Seeds usually flattened-concave, more rarely ovoid, ecarunculate, with endosperm Fruit a drupe, sometimes flattened, more rarely a capsule; mesocarp fleshy; endocarp woody, bony or stony, thin to thick and massive, usually tubercled, muricate, pitted or ridged, sometimes dehiscent by valves Ovary superior, 2–5-locular, with 1 (–2) apical pendulous orthotropous or anatropous ovules per locule, without obturator; styles 1–5, simple or bipartite Stamens 5, 10 or 15, uniseriate or biseriate, sometimes unequal; anthers introrse, 2-thecous, opening lengthwise; staminodes sometimes present in ? flowers; pistillode of ? flowers linear-subulate, cylindric or narrowly conical, sometimes pentagonal at base