The World Flora Online

Arbres ou arbustes, rarement herbes, sans latex, glabres ou parfois à poils simples. Stipules présentes, caduques ou persistantes, entières ou divisées. Feuilles alternes, simples, pétiolées ; limbe à marge entière à denticulée ou serretée ; nervation pennée. Inflorescences terminales ou axillaires, sous forme de racèmes, panicules, cymes, fascicules ou fleurs solitaires, l’unité ultime cymeuse ou racémeuse. Fleurs bisexuées, actinomorphes ou rarement zygomorphes ; pédicelles souvent articulés ; sépales (3–)5(–10), libres, caducs ou persistants, parfois accrescents en fruit ; pétales (4–)5(–10), libres, imbriqués ou contortés en bouton, souvent rapidement caducs ; étamines 1, 5 ou 10–∞, libres, anthères extrorses, basifixes, à déhiscence par pores apicaux ou parfois par fentes longitudinales, staminodes parfois présents ; disque absent ; ovaire supère, syncarpique mais parfois profondément divisé et carpelles presque libres en fruit, 1–5-loculaire, à placentation marginale et 1–∞ ovules par loge ; style 1, apical ou gynobasique, entier ou divisé au sommet. Fruits capsulaires à déhiscence septicide ou apocarpe avec drupéoles à 1 graine insérée sur le réceptacle accrescent. Graines non arillées, albuminées ou non, ailées ou non ; embryon droit ou courbé ; cotylédons droits ou courbés, incombants (l’un sur l’autre) ou accombants (l’un à côté de l’autre), égaux (graine isocotylée) ou inégaux (graine hétérocotylée).

Famille d’environ 32 genres et 500 espèces, dans toutes les régions tropicales, mais presque absente de l’Australie ; au Gabon, 7 genres et 41 espèces.

Arbres , arbustes, suffrutex ou herbes.'Feuilles'alternes, simples, rarement composées-pennées, généralement glabres, à nervures secondaires espacées ou très rapprochées; stipules entières ou divisées, caduques ou persistantes.'Inflorescences': racèmes, panicules, cymes, fascicules ou fleurs solitaires.'Fleurs'♀ actinomorphes, rarement zygomorphes; sépales (4)5(10), caducs ou persistants; pétales (4)5(10), caducs; étamines 1, 5, 8-12-, à filet persistant; anthères linéaires, parfois épaissies à la base, basifixes, à thèques lisses ou ridées transversalement, déhiscentes par fente longitudinale ou par pores apicaux; staminodes parfois présents; gynécée à carpelles libres ou soudés; ovaire supère; ovules 1-∞ par loge; style entier ou divisé au sommet.'Fruits'à drupéoles insérées sur le réceptacle ± accru (torus), baies, akènes ou capsules.'Graines albuminées ou exalbuminées, à embryon droit ou courbe, incombant ou accombant, isocotylé ou hétérocotylé.\n\t\t\tFamille comprenant 34 genres et environ 400 espèces dans les régions tropicales et subtropicales; pour la Flore, 6 genres, 45 espèces et 3 variétés.\n\t\t\tLes rameaux de certaines espèces des genres'Ochna ,'Campylospermum'et'Rhabdophyllum ont un mode de croissance qui semble saisonnier : alternance de parties normalement feuillées et de parties aphylles garnies uniquement d'écailles gemmaires persistantes.\n\t\t\tChez les'Ochna ,'Campylospermum ,'Rhabdophyllum'et'Idertia , les fleurs sont jaune vif; à maturité, les drupéoles ont une couleur noir brillant et sont entourées des sépales accrescents rouge vif. Les'Lophira'ont des fleurs blanches et les'Sauvagesia ont des fleurs blanchâtres, roses, rougeâtres ou violacées.

Petals free, 4–10, subsessile, contorted or imbricate Stamens few to many, free; staminodes sometimes present, subulate or petaloid, sometimes connate into a tube; filaments persistent; anthers linear, basifixed, opening lengthwise or by terminal pores Flowers hermaphrodite, actinomorphic, mostly racemose or paniculate Sepals 4–5, rarely 10, free, imbricate or rarely contorted Trees or shrubs with watery juice, rarely herbs Leaves alternate, simple, very rarely pinnate, often with numerous pinnate nerves; stipules present, sometimes laciniate Ovary entire to deeply lobed, 1–10-celled; ovules 1 to many, axile or parietal or attached to the intrusive placentas; style simple or divided at the apex Fruiting carpels often becoming quite separate on the enlarged torus and drupaceous, or elongated capsular and septicidal Seeds 1 to many, with or without endosperm; embryo usually straight Stamens few to many, free; staminodes sometimes present, subulate or petaloid, sometimes connate into a tube; filaments persistent; anthers linear, basifixed, opening lengthwise or by terminal pores Flowers hermaphrodite, actinomorphic, mostly racemose or paniculate Sepals 4–5, rarely 10, free, imbricate or rarely contorted Trees or shrubs with watery juice, rarely herbs Leaves alternate, simple, very rarely pinnate, often with numerous pinnate nerves; stipules present, sometimes laciniate Ovary entire to deeply lobed, 1–10-celled; ovules 1 to many, axile or parietal or attached to the intrusive placentas; style simple or divided at the apex Fruiting carpels often becoming quite separate on the enlarged torus and drupaceous, or elongated capsular and septicidal Seeds 1 to many, with or without endosperm; embryo usually straight

Leaves alternate, simple or very rarely pinnately compound, entire or more often with undulate to serrate or ciliate margins, penninerved (often densely so), membranous to coriaceous, with stipules entire to fringed or ± dissected, free or ± united intrapetiolarly, caducous or ± persistent Flowers bisexual, actinomorphic or rarely d: zygomorphic, sometimes fragrant, in terminal or axillary paniculate to umbellate or fascicled cymes or in racemes or solitary; pedicels usually articulated Trees, shrubs or shrublets, or more rarely perennial or annual herbs, glabrous or rarely shortly pubescent; bark rough or smooth, sometimes flaking, occasionally pigmented beneath; stem with cortical vascular bundles Petals (4) 5 (6–12), free, contorted in bud, deciduous Sepals (3–4) 5 (6–10), imbricate (usually quincuncial) in bud, free or rarely shortly united at the base, persistent or deciduous, sometimes enlarging in fruit Fruit either a septicidal 2–5-valved l-?-seeded capsule, or of (3) 4–12 1-seeded drupelets with fleshy or coriaceous mesocarp borne on a ± swollen receptacle Ovary free, sessile, syncarpous, either entire to shallowly lobed with apical style and 2–5 parietal (more rarely axile or basal) placentas each bearing 1-? ovules in two rows, or with (3) 5 (6–15) 1-ovulate lobes and gynobasic style; styles as many as the placentas or ovary-lobes, free towards the apex or completely united Seeds exarillate, with or without endosperm; embryo straight or curved, incumbent or accumbent, isocotylous or heterocotylous Stamens (1) 5-?, apparently whorled or diplos-temonous or antisepalous, more rarely excentrically grouped, free, inserted on the receptacle; staminodes sometimes present outside the stamens, subulate or petaloid, sometimes connate into a tube; anthers usually ± elongated, extrorse, basifixed, often deciduous, dehiscing by longitudinal slits or apical pores; connective occasionally with a slender prolongation; filaments persistent Flowers bisexual, actinomorphic or rarely d: zygomorphic, sometimes fragrant, in terminal or axillary paniculate to umbellate or fascicled cymes or in racemes or solitary; pedicels usually articulated Trees, shrubs or shrublets, or more rarely perennial or annual herbs, glabrous or rarely shortly pubescent; bark rough or smooth, sometimes flaking, occasionally pigmented beneath; stem with cortical vascular bundles Petals (4) 5 (6–12), free, contorted in bud, deciduous Sepals (3–4) 5 (6–10), imbricate (usually quincuncial) in bud, free or rarely shortly united at the base, persistent or deciduous, sometimes enlarging in fruit Fruit either a septicidal 2–5-valved l-?-seeded capsule, or of (3) 4–12 1-seeded drupelets with fleshy or coriaceous mesocarp borne on a ± swollen receptacle Ovary free, sessile, syncarpous, either entire to shallowly lobed with apical style and 2–5 parietal (more rarely axile or basal) placentas each bearing 1-? ovules in two rows, or with (3) 5 (6–15) 1-ovulate lobes and gynobasic style; styles as many as the placentas or ovary-lobes, free towards the apex or completely united Seeds exarillate, with or without endosperm; embryo straight or curved, incumbent or accumbent, isocotylous or heterocotylous Stamens (1) 5-?, apparently whorled or diplos-temonous or antisepalous, more rarely excentrically grouped, free, inserted on the receptacle; staminodes sometimes present outside the stamens, subulate or petaloid, sometimes connate into a tube; anthers usually ± elongated, extrorse, basifixed, often deciduous, dehiscing by longitudinal slits or apical pores; connective occasionally with a slender prolongation; filaments persistent

Seeds with or without endosperm Ovary superior, sessile, syncarpous, entire to lobed with style and 2–5 parietal placentas each with 1–many ovules, or with (3–)5(–6–15) lobes each with 1 ovule and style gynobasic; styles as many as placentas or ovary-lobes but completely united or free only at the apex with one globose or lobed stigma or separate stigmas Fruit a collection of 3–12 one-seeded drupelets borne on a fleshy enlarged receptacle, a nut surrounded by unequal enlarged sepals or a septicidal 2–5-valved, 1–many-seeded capsule Stamens few to many, with persistent filaments; anthers linear, basifixed, dehiscing lengthwise or by apical pores; staminodes present outside stamens in a few genera (>i>Sauvagesia and allies) Petals (4–)5(–6–12), free, contorted in bud, often clawed, deciduous Sepals (3–4–)5(–6–10), free, usually quincuncially imbricate, persistent or deciduous Flowers hermaphrodite, almost always regular, solitary or in fascicles or raceme-like, paniculate or cymose inflorescences; pedicels often articulated Trees, shrubs or geophytic shrublets, less often annual or perennial herbs Leaves alternate, simple (pinnate and opposite in Rhytidanthera, a small S American genus) often with numerous lateral veins and densely reticulate tertiary venation, entire to serrate or setulose; stipules entire to deeply divided, deciduous or persistent Ovary superior, sessile, syncarpous, entire to lobed with style and 2–5 parietal placentas each with 1–many ovules, or with (3–)5(–6–15) lobes each with 1 ovule and style gynobasic; styles as many as placentas or ovary-lobes but completely united or free only at the apex with one globose or lobed stigma or separate stigmas Fruit a collection of 3–12 one-seeded drupelets borne on a fleshy enlarged receptacle, a nut surrounded by unequal enlarged sepals or a septicidal 2–5-valved, 1–many-seeded capsule Stamens few to many, with persistent filaments; anthers linear, basifixed, dehiscing lengthwise or by apical pores; staminodes present outside stamens in a few genera (>i>Sauvagesia and allies) Petals (4–)5(–6–12), free, contorted in bud, often clawed, deciduous Sepals (3–4–)5(–6–10), free, usually quincuncially imbricate, persistent or deciduous Flowers hermaphrodite, almost always regular, solitary or in fascicles or raceme-like, paniculate or cymose inflorescences; pedicels often articulated Trees, shrubs or geophytic shrublets, less often annual or perennial herbs Leaves alternate, simple (pinnate and opposite in Rhytidanthera, a small S American genus) often with numerous lateral veins and densely reticulate tertiary venation, entire to serrate or setulose; stipules entire to deeply divided, deciduous or persistent

Trees and shrubs, rarely herbs. Leaves alternate, simple, rarely pinnately compound, petiolate; stipules entire or sometimes lacerate, persistent or caducous; leaf blade margin dentate, serrate or rarely entire; veins pinnate. Inflorescences terminal or axillary, cymose or racemose, rarely 1-flowered. Flowers generally bisexual, actinomorphic or more rarely zygomorphic, bracteate; pedicels articulated. Sepals (2-4)5(10-15), free or more rarely united, imbricate or valvate. Petals (3-)5(-8), mostly free, clawed to sessile, contorted or imbricate. Stamens 5-10 or numerous; filaments generally free and persistent, or anthers sessile; anthers basifixed, longitudinally dehiscent or poricidal; staminodes present or not, sometimes persistent, awl-shaped, spatulate, or petaloid, sometimes connected into a tube. Gynoecium (2 or)3-5(-15)-carpellate, gynophore present or rarely absent; ovary superior, entire or deeply lobed, generally long styled; placentation basal, axile or parietal, rarely laminar; ovules 1 or 2 or numerous per locule; stigma entire or sometimes shortly divided at apex. Fruit fleshy or non-fleshy, generally a septicidal capsule, rarely a nut with accrescent sepals or a drupe, or separating into up to 15 blackish drupelets on a colored accrescent receptacle. Seeds endospermic or non-endospermic, winged or not; germination phanerocotylar or cryptocotylar; embryo usually straight, more rarely curved.