Shrubs or small trees , evergreen or deciduous, usually aromatic and resinous. Roots commonly with nitrogen-fixing nodules. Leaves alternate, simple or pinnatifid; stipules absent or present; petiole present. Leaf blade commonly with peltate, multicellular, glandular trichomes. Inflorescences axillary catkins; bracts present. Flowers usually unisexual, occasionally bisexual, staminate and pistillate flowers usually on different plants, occasionally on same plants; perianth absent. Staminate flowers subtended by solitary bract; stamens 2-14(-22), hypogynous or ± epigynous; filaments filiform, distinct or basally connate; anthers dorsifixed, 2-locular, extrorsely dehiscent by longitudinal slits. Pistillate flowers subtended by solitary bract, bracteoles present or absent, usually 2-4(-8); pistils 1, 2-carpellate, 1-locular; ovules 1, basal, erect; styles, if present, short; stigmas 2. Fruits drupaceous or nutlike, smooth or often covered with warty protuberances, these commonly with waxy coating; fruits sometimes enclosed by persistent, accrescent bracts and bracteoles. Seeds with little or no endosperm; embryo straight, with 2 plano-convex cotyledons. Trees or shrubs, evergreen or deciduous, monoecious or dioecious, aromatic, often with resinous, peltate glands. Leaves alternate, simple, pinnately veined, entire to irregularly serrate or lobed, rarely pinnatifid; stipules absent or rarely present. Flowers in spikes, usually unisexual, anemophilous, without perianth. Male flowers solitary in axil of each bract, with 2-4 bracteoles or not. Stamens 2-20, often 4-8 on receptacle at base of bract; filaments short, free or slightly united at base; anthers erect, dithecal, extrorse, dehiscing longitudinally. Female flowers solitary or 2-4 in axil of bract, usually with 2-4 bracteoles. Gynoecium of 2 carpels united into a compound and 1-loculed ovary; styles distinct or united only at base; ovule solitary, basal, erect, orthotropous. Fruit drupaceous, or nearly a nutlet, often with headlike, wax-covered papillae; endocarp hard. Seeds nearly without endosperm; embryo straight; cotyledons fleshy, plano-convex. Fls tiny, borne in unisexual catkins, without perianth, commonly one fl (or pseudanthial partial infl) in the axil of each bract; staminate fls commonly (including our spp.) with 2–8 (often 4) stamens; ovary bicarpellate, unilocular, the 2 short styles distinct or united at base, each with a linear-elongate stigma; ovule solitary, basal, erect, orthotropous; fr an achene or small nut or a small, thin-skinned drupe, sometimes enveloped by persistent, accrescent bracteoles; seeds with a straight, dicotyledonous embryo, nearly or quite without endosperm; aromatic, anemophilous shrubs or small trees with alternate, simple, pinnately veined lvs that characteristically bear 2 sorts of trichomes, one sort elongate, white, slender, and unicellular, the other sort multicellular, with a short, ± embedded stalk and an expanded, glandular or resinous, yellow or brownish head. 3/50. Sepals and petals absent, or the female with a few sepal-like whorled bracteoles Male flower subtended by a solitary bract: stamens 2 or more; anthers 2-celled Leaves alternate, simple, sometimes pinnately lobed, scaly-glandular beneath, exstipulate Flowers unisexual, monoecious or dioecious, in axillary spikes Fruit a drupe, often warted, the warts waxy Seed erect, without endosperm; embryo straight Female flower: ovary sessile, 1-celled; style short, 2-branched Ovule 1, erect, basal Trees or shrubs, often aromatic Leaves alternate, entire to irregularly dentate or lobed, occasionally pinnatifid, usually with aromatic resinous gland-dots; stipules usually absent Monoecious or dioecious trees, shrubs or subshrubs; nitrogen-fixing root-nodules present Seed with membranous testa and little or no endosperm Fruit either dry and with attached wing-like accrescent bracteoles or a small globose or ovoid drupe densely covered with bead-like often wax-covered papillae; endocarp hard Female flowers: ovary sessile, 1-locular; style short with 2 slender branches; ovule 1, erect from the base, orthotropous Male flowers: stamens 2–10(22); filaments short, free or partially united into a column; anthers erect, 2-thecous, opening lengthwise Perianth absent except in >i>Canacomyrica Flowers in dense elongate or globose axillary spikes (catkins), unisexual (male sometimes with vestigial female parts or some morphologically bisexual in Canacomyrica); bracts present and sometimes also smaller bracteoles Seed with membranous testa and little or no endosperm Female flowers: ovary sessile, 1-locular; style short with 2 slender branches; ovule 1, erect from the base, orthotropous Fruit either dry and with attached wing-like accrescent bracteoles or a small globose or ovoid drupe densely covered with bead-like often wax-covered papillae; endocarp hard Flowers in dense elongate or globose axillary spikes (catkins), unisexual (or some morphologically bisexual in Canacomyrica); bracts present and sometimes a pair of smaller bracteoles; perianth absent save in Canacomyrica Male flowers: stamens 2–10(–22); filaments short, free or partially united into a column; anthers erect, 2-thecous, opening lengthwise Leaves alternate, entire to irregularly dentate or lobed, occasionally pinnatifid, usually with aromatic resinous gland-dots Stipules usually absent Monoecious or dioecious trees, shrubs or subshrubs; nitrogen-fixing root-nodules present Arbustes'ou arbres, dioïques, plus rarement monoïques.'Feuilles'alternes, dépourvues de stipules, penninerves, à bords entiers, ondulés ou dentés, généralement persistantes et coriaces, parsemées de glandes surtout en dessous et aromatiques.'Épis'axillaires, solitaires, fasciculés, réunis en glomérules ou en épis composés.'Fleurs'♂ ♀, nues, insérées à l'aisselle d'une bractée. Les ♂ à 1-12 étamines; anthères 2-loculaires à déhiscence longitudinale. Les ♀ à ovaire 1-loculaire, muni à la base de bractéoles1; ovule 1, orthotrope, unitégumenté; style court divisé en deux branches stigmatiques lancéolées ou linéaires à filiformes.'Fruits'drupacés, ± globuleux, verruqueux et glandulaires, souvent recouverts d'un enduit cireux.'Graines à testa membraneux, dépourvues d'albumen; embryon droit.\n\t\t\tUn genre, bien représenté à l'époque tertiaire et groupant actuellement ± 70 espèces réparties dans les régions tempérées et les régions tropicales surtout de haute altitude. SELECTED REFERENCES Baird, J. R. 1968. A Taxonomic Revision of the Plant Family Myricaceae of North America, North of Mexico. Ph.D. thesis. University of North Carolina. Chevalier, A. 1901. Monographie des Myricacées. Mém. Soc. Sci. Nat. Cherbourg 32: 85-340. Elias, T. S. 1971. The genera of Myricaceae in the southeastern United States. J. Arnold Arbor. 52: 305-318. Sheffy, M. V. 1972. A Study of the Myricaceae from Eocene Sediments of Southeastern North America. Ph.D. thesis. Indiana University. Wilbur, R. L. 1994. The Myricaceae of the United States and Canada: Genera, subgenera, and series. Sida 16(1): 93-107. Youngken, H. W. 1919. The comparative morphology, taxonomy and distribution of the Myricaceae of the eastern United States. Contr. Bot. Lab. Morris Arbor. Univ. Pennsylvania 4: 339-400. SELECTED REFERENCES Baird, J. R. 1968. A Taxonomic Revision of the Plant Family Myricaceae of North America, North of Mexico. Ph.D. thesis. University of North Carolina. Chevalier, A. 1901. Monographie des Myricacées. Mém. Soc. Sci. Nat. Cherbourg 32: 85-340. Elias, T. S. 1971. The genera of Myricaceae in the southeastern United States. J. Arnold Arbor. 52: 305-318. Sheffy, M. V. 1972. A Study of the Myricaceae from Eocene Sediments of Southeastern North America. Ph.D. thesis. Indiana University. Wilbur, R. L. 1994. The Myricaceae of the United States and Canada: Genera, subgenera, and series. Sida 16(1): 93-107. Youngken, H. W. 1919. The comparative morphology, taxonomy and distribution of the Myricaceae of the eastern United States. Contr. Bot. Lab. Morris Arbor. Univ. Pennsylvania 4: 339-400. Shrubs or small trees , evergreen or deciduous, usually aromatic and resinous. Roots commonly with nitrogen-fixing nodules. Leaves alternate, simple or pinnatifid; stipules absent or present; petiole present. Leaf blade commonly with peltate, multicellular, glandular trichomes. Inflorescences axillary catkins; bracts present. Flowers usually unisexual, occasionally bisexual, staminate and pistillate flowers usually on different plants, occasionally on same plants; perianth absent. Staminate flowers subtended by solitary bract; stamens 2-14(-22), hypogynous or ± epigynous; filaments filiform, distinct or basally connate; anthers dorsifixed, 2-locular, extrorsely dehiscent by longitudinal slits. Pistillate flowers subtended by solitary bract, bracteoles present or absent, usually 2-4(-8); pistils 1, 2-carpellate, 1-locular; ovules 1, basal, erect; styles, if present, short; stigmas 2. Fruits drupaceous or nutlike, smooth or often covered with warty protuberances, these commonly with waxy coating; fruits sometimes enclosed by persistent, accrescent bracts and bracteoles. Seeds with little or no endosperm; embryo straight, with 2 plano-convex cotyledons. Arbustes'ou arbres, dioïques, plus rarement monoïques.'Feuilles'alternes, dépourvues de stipules, penninerves, à bords entiers, ondulés ou dentés, généralement persistantes et coriaces, parsemées de glandes surtout en dessous et aromatiques.'Épis'axillaires, solitaires, fasciculés, réunis en glomérules ou en épis composés.'Fleurs'♂ ♀, nues, insérées à l'aisselle d'une bractée. Les ♂ à 1-12 étamines; anthères 2-loculaires à déhiscence longitudinale. Les ♀ à ovaire 1-loculaire, muni à la base de bractéoles1; ovule 1, orthotrope, unitégumenté; style court divisé en deux branches stigmatiques lancéolées ou linéaires à filiformes.'Fruits'drupacés, ± globuleux, verruqueux et glandulaires, souvent recouverts d'un enduit cireux.'Graines à testa membraneux, dépourvues d'albumen; embryon droit.\n\t\t\tUn genre, bien représenté à l'époque tertiaire et groupant actuellement ± 70 espèces réparties dans les régions tempérées et les régions tropicales surtout de haute altitude. Sepals and petals absent, or the female with a few sepal-like whorled bracteoles Male flower subtended by a solitary bract: stamens 2 or more; anthers 2-celled Leaves alternate, simple, sometimes pinnately lobed, scaly-glandular beneath, exstipulate Flowers unisexual, monoecious or dioecious, in axillary spikes Fruit a drupe, often warted, the warts waxy Seed erect, without endosperm; embryo straight Female flower: ovary sessile, 1-celled; style short, 2-branched Ovule 1, erect, basal Trees or shrubs, often aromatic Male flower subtended by a solitary bract: stamens 2 or more; anthers 2-celled Leaves alternate, simple, sometimes pinnately lobed, scaly-glandular beneath, exstipulate Flowers unisexual, monoecious or dioecious, in axillary spikes Fruit a drupe, often warted, the warts waxy Seed erect, without endosperm; embryo straight Female flower: ovary sessile, 1-celled; style short, 2-branched Ovule 1, erect, basal Trees or shrubs, often aromatic Leaves alternate, entire to irregularly dentate or lobed, occasionally pinnatifid, usually with aromatic resinous gland-dots; stipules usually absent Monoecious or dioecious trees, shrubs or subshrubs; nitrogen-fixing root-nodules present Seed with membranous testa and little or no endosperm Fruit either dry and with attached wing-like accrescent bracteoles or a small globose or ovoid drupe densely covered with bead-like often wax-covered papillae; endocarp hard Female flowers: ovary sessile, 1-locular; style short with 2 slender branches; ovule 1, erect from the base, orthotropous Male flowers: stamens 2–10(22); filaments short, free or partially united into a column; anthers erect, 2-thecous, opening lengthwise Perianth absent except in >i>Canacomyrica Flowers in dense elongate or globose axillary spikes (catkins), unisexual (male sometimes with vestigial female parts or some morphologically bisexual in Canacomyrica); bracts present and sometimes also smaller bracteoles Monoecious or dioecious trees, shrubs or subshrubs; nitrogen-fixing root-nodules present Seed with membranous testa and little or no endosperm Fruit either dry and with attached wing-like accrescent bracteoles or a small globose or ovoid drupe densely covered with bead-like often wax-covered papillae; endocarp hard Female flowers: ovary sessile, 1-locular; style short with 2 slender branches; ovule 1, erect from the base, orthotropous Male flowers: stamens 2–10(22); filaments short, free or partially united into a column; anthers erect, 2-thecous, opening lengthwise Perianth absent except in >i>Canacomyrica Flowers in dense elongate or globose axillary spikes (catkins), unisexual (male sometimes with vestigial female parts or some morphologically bisexual in Canacomyrica); bracts present and sometimes also smaller bracteoles Fls tiny, borne in unisexual catkins, without perianth, commonly one fl (or pseudanthial partial infl) in the axil of each bract; staminate fls commonly (including our spp.) with 2–8 (often 4) stamens; ovary bicarpellate, unilocular, the 2 short styles distinct or united at base, each with a linear-elongate stigma; ovule solitary, basal, erect, orthotropous; fr an achene or small nut or a small, thin-skinned drupe, sometimes enveloped by persistent, accrescent bracteoles; seeds with a straight, dicotyledonous embryo, nearly or quite without endosperm; aromatic, anemophilous shrubs or small trees with alternate, simple, pinnately veined lvs that characteristically bear 2 sorts of trichomes, one sort elongate, white, slender, and unicellular, the other sort multicellular, with a short, ± embedded stalk and an expanded, glandular or resinous, yellow or brownish head. 3/50. Trees or shrubs, evergreen or deciduous, monoecious or dioecious, aromatic, often with resinous, peltate glands. Leaves alternate, simple, pinnately veined, entire to irregularly serrate or lobed, rarely pinnatifid; stipules absent or rarely present. Flowers in spikes, usually unisexual, anemophilous, without perianth. Male flowers solitary in axil of each bract, with 2-4 bracteoles or not. Stamens 2-20, often 4-8 on receptacle at base of bract; filaments short, free or slightly united at base; anthers erect, dithecal, extrorse, dehiscing longitudinally. Female flowers solitary or 2-4 in axil of bract, usually with 2-4 bracteoles. Gynoecium of 2 carpels united into a compound and 1-loculed ovary; styles distinct or united only at base; ovule solitary, basal, erect, orthotropous. Fruit drupaceous, or nearly a nutlet, often with headlike, wax-covered papillae; endocarp hard. Seeds nearly without endosperm; embryo straight; cotyledons fleshy, plano-convex. Seed with membranous testa and little or no endosperm Female flowers: ovary sessile, 1-locular; style short with 2 slender branches; ovule 1, erect from the base, orthotropous Fruit either dry and with attached wing-like accrescent bracteoles or a small globose or ovoid drupe densely covered with bead-like often wax-covered papillae; endocarp hard Flowers in dense elongate or globose axillary spikes (catkins), unisexual (or some morphologically bisexual in Canacomyrica); bracts present and sometimes a pair of smaller bracteoles; perianth absent save in Canacomyrica Male flowers: stamens 2–10(–22); filaments short, free or partially united into a column; anthers erect, 2-thecous, opening lengthwise Leaves alternate, entire to irregularly dentate or lobed, occasionally pinnatifid, usually with aromatic resinous gland-dots Stipules usually absent Monoecious or dioecious trees, shrubs or subshrubs; nitrogen-fixing root-nodules present Female flowers: ovary sessile, 1-locular; style short with 2 slender branches; ovule 1, erect from the base, orthotropous Fruit either dry and with attached wing-like accrescent bracteoles or a small globose or ovoid drupe densely covered with bead-like often wax-covered papillae; endocarp hard Flowers in dense elongate or globose axillary spikes (catkins), unisexual (or some morphologically bisexual in Canacomyrica); bracts present and sometimes a pair of smaller bracteoles; perianth absent save in Canacomyrica Male flowers: stamens 2–10(–22); filaments short, free or partially united into a column; anthers erect, 2-thecous, opening lengthwise Leaves alternate, entire to irregularly dentate or lobed, occasionally pinnatifid, usually with aromatic resinous gland-dots Stipules usually absent Monoecious or dioecious trees, shrubs or subshrubs; nitrogen-fixing root-nodules presentGeneral Information
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Morphology
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Literature
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Flora of North America @ efloras.org
LiteratureFlore d'Afrique Centrale
MorphologyFlora of West Tropical Africa - species descriptions
MorphologyFlora Zambesiaca - descriptions
MorphologyNortheastern Flora
General InformationFlora of China @ efloras.org
General InformationPlants Of the World Online Portal - FTEA
Morphology
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Wax-myrtle Family |
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