Shrubs or trees. Stipules intrapetiolar. Leaves alternate or rarely opposite, simple; leaf blade margin entire. Flowers usually bisexual, in axillary fascicles or cymes, regular, 5-merous, often heterostylous. Sepals 5, basally connate, with imbricate or valvate lobes, persistent. Petals 5, distinct, imbricate, usually with a scale on inner face at base. Stamens 5, 10, or 20, 1- or 2-verticillate; filament bases usually connate into a tube; anthers elliptic, 2-celled, with longitudinal slits. Ovary superior, connected with 3-5 carpels, 3-5-locular, with 1(or 2) axile; ovules pendulous, anatropous to hemitropous, placentation axile; styles 1-3 or 5, distinct or somewhat connate; stigmas oblique. Fruit a capsule or a 1-seeded drupe. Seeds with straight embryo and copious (rarely absent) endosperm. Calyx persistent, campanulate, lobes 5, imbricate Flowers fasciculate, hermaphrodite, rarely subdioecious, hypogynous, actinomorphic Leaves alternate, rarely opposite, simple, entire; stipules intrapetiolar, rarely extrapetiolar, often caducous Trees, shrubs, or undershrubs Fruit drupaceous; seeds with or without endosperm; embryo straight Ovary of 3 carpels, 3-celled, mostly two of the cells sterile, fertile cells 1–2-ovuled; ovules pendulous; styles 3, free or more or less connate; stigmas oblique, depressed-capitate or clavate Petals 5, free, deciduous, imbricate, mostly ligulate on the inside Stamens 10, in 2 series, more or less connate at the base; anthers ellipsoid, 2-celled, opening lengthwise Fruit a 1-seeded fleshy drupe or rarely a 3(–4)-locular, 3(–4)-seeded capsule dehiscing longitudinally Seeds with no aril and no or very little endosperm; embryo straight with flat or semiconvex cotyledons Stamens 10, all fertile, the filaments united at the base to form a shallow or urn-shaped cup, sometimes glandular; anthers basifixed, dehiscing longitudinally Ovary superior, 2–3(–4)-locular, the locules with 1 or rarely 2 pendulous ovules; styles 2–3(–4), free or ± united, with club-shaped, capitate or rarely acute stigmas Sepals (4–)5, valvate, ± united Petals (4–)5, with contorted aestivation, free, clawed, soon falling, usually with a ligulate nectariferous appendage inside Leaves alternate or rarely opposite, simple, entire, pinnately nerved; stipules ± united, intrapetiolar or rarely interpetiolar Flowers axillary, solitary, in fascicles or rarely in pedunculate inflorescences, regular, hermaphrodite or unisexual, heterostylous Trees, shrubs or subshrubs, glabrous Fruit a 1-seeded fleshy drupe (or rarely a 3 (4)-locular 3 (4)-seeded capsule dehiscing longitudinally) Seeds not compressed, exarillate, without or with scanty endosperm; embryo straight, with flat or semiconvex cotyledons Trees, shrubs or shrublets, glabrous Leaves alternate (or rarely opposite), simple, entire, penninerved, with stipules ± united and intrapetiolar (or rarely interpetiolar) Ovary superior, 2–3 (4)-locular, with each loculus 1-ovulate (or rarely 2-ovulate); ovules pendulous; styles 2–3 (4), free or ± united, with stigmas clavate to obliquely capitate or depressed-capitate rarely acute Stamens 5 + 5, all fertile, with filaments united at the base to form a deep urceole or shallow rarely glandiferous cup; anthers basifixed, dehiscing longitudinally Petals (4) 5, contorted in bud, free, unguiculate, caducous, usually with a ventral ligule-like nectariferous appendage Sepals (4) 5, valvate, ± united Flowers axillary, solitary or in fascicles (rarely pedunculate), actinomorphic, bisexual or rarely subdioecious, heterostylic Arbres ou arbrisseaux glabres. Feuilles alternes ou opposées, simples, entières, penninerves, pétiolées; stipules intrapétiolaires ± soudées, rarement stipules extrapétiolaires. Fleurs axillaires, solitaires ou fasciculées, hermaphrodites, actinomorphes, hétérostylées. Sépales 5, soudés à la base. Pétales 5, libres, contortés, munis à la face ventrale d'un appendice nectarifère. Étamines 10, à filets unis à la base en un tube; anthères basifixes, à déhiscence longitudinale. Ovaire supère, 2-3(-4)-loculaire à 1 (rarement 2) ovule pendant par loge. Styles 3, libres ou ± soudés; stigmates 3. Drupes monospermes ou capsules 3 (-4)-loculaires, à 1 graine par loge, à déhiscence longitudinale. Arbres , arbrisseaux.'Feuilles'alternes, distiques, entières; stipules intrapétiolaires, parfois soudées en une pièce triangulaire ± longuement persistante.'Inflorescences'en fascicules, racèmes fasciculiformes, parfois 1-flores.'Fleurs'☿; calice à 5 segments ± dilatés à la base; corolle à 5 pétales libres, onguiculés, munis à la base, en dedans, d'un appendice squamiforme, en général plus longs que le calice; étamines 10, obdiplostémones, en 2 verticilles; filets ± connés à la base; anthères basifixes, à déhiscence longitudinale; ovaire 2-3-loculaire, à 1 ovule penduleux par loge; styles ± longuement soudés.'Fruits'drupacés, peu charnus, à 1 graine.'Graines sans albumen ou à albumen ± réduit.\n\t\t\tFamille comprenant 2 genres et ± 200 espèces; au Congo belge, 2 genres et 3 espèces. Arbres ou arbrisseaux glabres. Feuilles alternes ou opposées, simples, entières, penninerves, pétiolées; stipules intrapétiolaires ± soudées, rarement stipules extrapétiolaires. Fleurs axillaires, solitaires ou fasciculées, hermaphrodites, actinomorphes, hétérostylées. Sépales 5, soudés à la base. Pétales 5, libres, contortés, munis à la face ventrale d'un appendice nectarifère. Étamines 10, à filets unis à la base en un tube; anthères basifixes, à déhiscence longitudinale. Ovaire supère, 2-3(-4)-loculaire à 1 (rarement 2) ovule pendant par loge. Styles 3, libres ou ± soudés; stigmates 3. Drupes monospermes ou capsules 3 (-4)-loculaires, à 1 graine par loge, à déhiscence longitudinale. Arbres , arbrisseaux.'Feuilles'alternes, distiques, entières; stipules intrapétiolaires, parfois soudées en une pièce triangulaire ± longuement persistante.'Inflorescences'en fascicules, racèmes fasciculiformes, parfois 1-flores.'Fleurs'☿; calice à 5 segments ± dilatés à la base; corolle à 5 pétales libres, onguiculés, munis à la base, en dedans, d'un appendice squamiforme, en général plus longs que le calice; étamines 10, obdiplostémones, en 2 verticilles; filets ± connés à la base; anthères basifixes, à déhiscence longitudinale; ovaire 2-3-loculaire, à 1 ovule penduleux par loge; styles ± longuement soudés.'Fruits'drupacés, peu charnus, à 1 graine.'Graines sans albumen ou à albumen ± réduit.\n\t\t\tFamille comprenant 2 genres et ± 200 espèces; au Congo belge, 2 genres et 3 espèces. Calyx persistent, campanulate, lobes 5, imbricate Flowers fasciculate, hermaphrodite, rarely subdioecious, hypogynous, actinomorphic Leaves alternate, rarely opposite, simple, entire; stipules intrapetiolar, rarely extrapetiolar, often caducous Trees, shrubs, or undershrubs Fruit drupaceous; seeds with or without endosperm; embryo straight Ovary of 3 carpels, 3-celled, mostly two of the cells sterile, fertile cells 1–2-ovuled; ovules pendulous; styles 3, free or more or less connate; stigmas oblique, depressed-capitate or clavate Petals 5, free, deciduous, imbricate, mostly ligulate on the inside Stamens 10, in 2 series, more or less connate at the base; anthers ellipsoid, 2-celled, opening lengthwise Flowers fasciculate, hermaphrodite, rarely subdioecious, hypogynous, actinomorphic Leaves alternate, rarely opposite, simple, entire; stipules intrapetiolar, rarely extrapetiolar, often caducous Trees, shrubs, or undershrubs Fruit drupaceous; seeds with or without endosperm; embryo straight Ovary of 3 carpels, 3-celled, mostly two of the cells sterile, fertile cells 1–2-ovuled; ovules pendulous; styles 3, free or more or less connate; stigmas oblique, depressed-capitate or clavate Petals 5, free, deciduous, imbricate, mostly ligulate on the inside Stamens 10, in 2 series, more or less connate at the base; anthers ellipsoid, 2-celled, opening lengthwise Fruit a 1-seeded fleshy drupe (or rarely a 3 (4)-locular 3 (4)-seeded capsule dehiscing longitudinally) Seeds not compressed, exarillate, without or with scanty endosperm; embryo straight, with flat or semiconvex cotyledons Trees, shrubs or shrublets, glabrous Leaves alternate (or rarely opposite), simple, entire, penninerved, with stipules ± united and intrapetiolar (or rarely interpetiolar) Ovary superior, 2–3 (4)-locular, with each loculus 1-ovulate (or rarely 2-ovulate); ovules pendulous; styles 2–3 (4), free or ± united, with stigmas clavate to obliquely capitate or depressed-capitate rarely acute Stamens 5 + 5, all fertile, with filaments united at the base to form a deep urceole or shallow rarely glandiferous cup; anthers basifixed, dehiscing longitudinally Petals (4) 5, contorted in bud, free, unguiculate, caducous, usually with a ventral ligule-like nectariferous appendage Sepals (4) 5, valvate, ± united Flowers axillary, solitary or in fascicles (rarely pedunculate), actinomorphic, bisexual or rarely subdioecious, heterostylic Seeds not compressed, exarillate, without or with scanty endosperm; embryo straight, with flat or semiconvex cotyledons Trees, shrubs or shrublets, glabrous Leaves alternate (or rarely opposite), simple, entire, penninerved, with stipules ± united and intrapetiolar (or rarely interpetiolar) Ovary superior, 2–3 (4)-locular, with each loculus 1-ovulate (or rarely 2-ovulate); ovules pendulous; styles 2–3 (4), free or ± united, with stigmas clavate to obliquely capitate or depressed-capitate rarely acute Stamens 5 + 5, all fertile, with filaments united at the base to form a deep urceole or shallow rarely glandiferous cup; anthers basifixed, dehiscing longitudinally Petals (4) 5, contorted in bud, free, unguiculate, caducous, usually with a ventral ligule-like nectariferous appendage Sepals (4) 5, valvate, ± united Flowers axillary, solitary or in fascicles (rarely pedunculate), actinomorphic, bisexual or rarely subdioecious, heterostylic Shrubs or trees. Stipules intrapetiolar. Leaves alternate or rarely opposite, simple; leaf blade margin entire. Flowers usually bisexual, in axillary fascicles or cymes, regular, 5-merous, often heterostylous. Sepals 5, basally connate, with imbricate or valvate lobes, persistent. Petals 5, distinct, imbricate, usually with a scale on inner face at base. Stamens 5, 10, or 20, 1- or 2-verticillate; filament bases usually connate into a tube; anthers elliptic, 2-celled, with longitudinal slits. Ovary superior, connected with 3-5 carpels, 3-5-locular, with 1(or 2) axile; ovules pendulous, anatropous to hemitropous, placentation axile; styles 1-3 or 5, distinct or somewhat connate; stigmas oblique. Fruit a capsule or a 1-seeded drupe. Seeds with straight embryo and copious (rarely absent) endosperm. Fruit a 1-seeded fleshy drupe or rarely a 3(–4)-locular, 3(–4)-seeded capsule dehiscing longitudinally Seeds with no aril and no or very little endosperm; embryo straight with flat or semiconvex cotyledons Stamens 10, all fertile, the filaments united at the base to form a shallow or urn-shaped cup, sometimes glandular; anthers basifixed, dehiscing longitudinally Ovary superior, 2–3(–4)-locular, the locules with 1 or rarely 2 pendulous ovules; styles 2–3(–4), free or ± united, with club-shaped, capitate or rarely acute stigmas Sepals (4–)5, valvate, ± united Petals (4–)5, with contorted aestivation, free, clawed, soon falling, usually with a ligulate nectariferous appendage inside Leaves alternate or rarely opposite, simple, entire, pinnately nerved; stipules ± united, intrapetiolar or rarely interpetiolar Flowers axillary, solitary, in fascicles or rarely in pedunculate inflorescences, regular, hermaphrodite or unisexual, heterostylous Trees, shrubs or subshrubs, glabrous Seeds with no aril and no or very little endosperm; embryo straight with flat or semiconvex cotyledons Stamens 10, all fertile, the filaments united at the base to form a shallow or urn-shaped cup, sometimes glandular; anthers basifixed, dehiscing longitudinally Ovary superior, 2–3(–4)-locular, the locules with 1 or rarely 2 pendulous ovules; styles 2–3(–4), free or ± united, with club-shaped, capitate or rarely acute stigmas Sepals (4–)5, valvate, ± united Petals (4–)5, with contorted aestivation, free, clawed, soon falling, usually with a ligulate nectariferous appendage inside Leaves alternate or rarely opposite, simple, entire, pinnately nerved; stipules ± united, intrapetiolar or rarely interpetiolar Flowers axillary, solitary, in fascicles or rarely in pedunculate inflorescences, regular, hermaphrodite or unisexual, heterostylous Trees, shrubs or subshrubs, glabrousGeneral Information
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Morphology
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