The World Flora Online

Arbres ou arbustes, résineux, souvent avec contreforts; indument à poils simples ou fasciculés. Feuilles alternes, stipulées, à pétiole géniculé ou non, simples, à marge entière ou ondulée, portant une glande à la base de la face supérieure; nervures secondaires pennées, nervation tertiaire scalariforme ou réticulée. Inflorescence axillaire ou terminale, en épi ou panicule; bractées présentes. Fleurs régulières, hermaphrodites; sépales 5, imbriqués ou valvaires, parfois soudés à la base; pétales 5, tordus en bouton, libres ou soudés à la base; étamines 5-110, à filets libres, anthères à 2 loges, basiversatile (en Afrique), base profondément cordée, connectif souvent prolongé; ovaire supère ou semi-infère, avec ou sans androgynophore, (2-)3-4(-5)-loculaire, parfois partiellement uniloculaire, à 1-4 ovules par loge; style 1, parfois divisé en trois au sommet. Fruits monospermes par avortement, généralement indéhiscents, libres ou inclus dans le calice accrescent qui forme des ailes. Graine généralement exalbuminée, cotylédons inégaux.

Seeds without endosperm; cotyledons thick and equal or variously contorted Fruit free from or included in the enlarged calyx-tube, rarely closely adnate to it, indehiscent, usually 1-seeded Ovary inserted on the broad torus or slightly immersed in it, 3-celled; style subulate or variously thickened, entire or 3-lobed; ovules 2 in each cell Stamens rarely definite, hypogynous or subperigynous; filaments short, often dilated at the base; anthers 2-celled, opening lengthwise; connective mucronate or aristate or club-shaped Petals 5, much contorted, free or slightly connate at the base, often hairy and inconspicuous Trees with resinous wood Leaves alternate, entire, penninerved; indumentum often of two kinds, of stellate hairs or rarely of peltáte scales; stipules mostly small and deciduous, sometimes very large and enclosing the young bud, rarely persistent, leaving an annular scar after falling away Flowers hermaphrodite, actinomorphic, fragrant, in axillary panicles Calyx-tube short or well developed, free or adnate to the ovary; lobes 5, imbricate or valvate, mostly greatly enlarged, wing-like, and ascending in fruit and then strongly nervose, rarely small and reflexed or spreading Fruit free from or included in the enlarged calyx-tube, rarely closely adnate to it, indehiscent, usually 1-seeded Ovary inserted on the broad torus or slightly immersed in it, 3-celled; style subulate or variously thickened, entire or 3-lobed; ovules 2 in each cell Stamens rarely definite, hypogynous or subperigynous; filaments short, often dilated at the base; anthers 2-celled, opening lengthwise; connective mucronate or aristate or club-shaped Petals 5, much contorted, free or slightly connate at the base, often hairy and inconspicuous Trees with resinous wood Leaves alternate, entire, penninerved; indumentum often of two kinds, of stellate hairs or rarely of peltáte scales; stipules mostly small and deciduous, sometimes very large and enclosing the young bud, rarely persistent, leaving an annular scar after falling away Flowers hermaphrodite, actinomorphic, fragrant, in axillary panicles Calyx-tube short or well developed, free or adnate to the ovary; lobes 5, imbricate or valvate, mostly greatly enlarged, wing-like, and ascending in fruit and then strongly nervose, rarely small and reflexed or spreading

Fruit 1-seeded, indehiscent, surrounded by the wings (always 5 in the African species) of the accrescent calyx Flowers actinomorphic, bisexual Leaves alternate, petiolate, entire, penninerved, in African species always with an extra-floral nectary at the base of the midrib on the upper face of the lamina; stipules caducous Trees or shrubs Ovary superior, (1) 3-locular with axile or parietal placentation; carpels (1) 3, 2-ovulate Stamens 5, 10, 15 or numerous (always numerous in African species) Petals 5, contorted to the right, free or slightly connate at the base Sepals 5, accrescent Flowers actinomorphic, bisexual Leaves alternate, petiolate, entire, penninerved, in African species always with an extra-floral nectary at the base of the midrib on the upper face of the lamina; stipules caducous Trees or shrubs Ovary superior, (1) 3-locular with axile or parietal placentation; carpels (1) 3, 2-ovulate Stamens 5, 10, 15 or numerous (always numerous in African species) Petals 5, contorted to the right, free or slightly connate at the base Sepals 5, accrescent

Trees, evergreen or semievergreen, rarely deciduous in dry season. Xylem with aromatic resin in intercellular resin canals. Branchlets with stipular scars, sometimes annular. Leaves simple, alternate; stipules persistent or caducous, large or small; leaf blade with lateral veins pinnate, margin entire or sinuate-crenate. Inflorescences few- or many-flowered, terminal or axillary racemes or panicles; flowers usually sweetly scented; bracts usually fugacious and minute, rarely persistent and large. Inflorescences, calyces, petals, ovary, and other parts usually with stellate, squamate, fascicled or free-standing hairs. Flowers bisexual, actinomorphic, contorted. Calyx lobes 5, free or united at base, imbricate in bud if not united. Petals 5, adnate or connate at base. Stamens (10-)15 to many, free from or connate to petals; filaments usually dilated at base; anthers 2-celled, with 2 pollen sacs per cell (Chinese species); connective appendages aristate, filiform or stout. Ovary superior, rarely semi-inferior, slightly immersed in torus, usually 3-loculed, each locule 2-, rarely many ovuled; ovules pendulous, lateral or anatropous. Fruit usually nutlike, sometimes capsular and 3-valved, 1(to many)-seeded, with persistent, variously accrescent calyx of which 2 or more lobes are usually developed into lorate wings. Seed exalbuminous; cotyledons fleshy, equal or unequal, applanate or ± folded or cerebriform, entire or laciniate; radicle directed toward hilum, usually included between cotyledons.

Small to large usually resinous mostly buttressed trees; indumentum various but often of fasciculate hairs, frequently glandular Stamens 5 to over 100; filaments free or connate; anthers 2-thecous, introrse or laterally dehiscent Ovary superior or semi-inferior, partly unilocular or (2–)3–5-locular (4 in the sole S. American genus); ovules 2(–4) in each locule, axile or parietal Fruit indehiscent, usually 1-seeded with woody pericarp, usually with 2–5 persistent wing-like sepals but not accrescent in a few genera Leaves alternate, simple, entire or sinuate at margin; stipules paired, large or small, persistent or soon falling Flowers ?, regular, in axillary or terminal branched panicles, racemose or less often cymose; bracts and bracteoles paired, persistent or soon falling Calyx persistent; lobes imbricate or valvate, free or fused at the base; 2–5 lobes usually accrescent into very distinct wings in fruiting state Corolla-lobes 5, contorted, connate at base or free Stamens 5 to over 100; filaments free or connate; anthers 2-thecous, introrse or laterally dehiscent Ovary superior or semi-inferior, partly unilocular or (2–)3–5-locular (4 in the sole S. American genus); ovules 2(–4) in each locule, axile or parietal Fruit indehiscent, usually 1-seeded with woody pericarp, usually with 2–5 persistent wing-like sepals but not accrescent in a few genera Leaves alternate, simple, entire or sinuate at margin; stipules paired, large or small, persistent or soon falling Flowers ?, regular, in axillary or terminal branched panicles, racemose or less often cymose; bracts and bracteoles paired, persistent or soon falling Calyx persistent; lobes imbricate or valvate, free or fused at the base; 2–5 lobes usually accrescent into very distinct wings in fruiting state Corolla-lobes 5, contorted, connate at base or free