Cratoneuron filicinum (Hedw.) Spruce
  • Cat. Musc.: 21. 1867. 


Cite taxon page as 'WFO (2025): Cratoneuron filicinum (Hedw.) Spruce. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-0001180980. Accessed on: 04 Jun 2025'

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General Information

Plants somewhat stiff. Stems with shoot and branch apices often pale; paraphyllia sometimes present only in youngest shoot portions; axillary hairs few, weak. Stem leaves gradually or suddenly narrowed to apex, plane or slightly concave; alar cells usually many, region large, usually strongly widened marginally, reaching from margin to costa or almost so; medial laminal cells 12-55 × 4-11 µm. Branch leaves usually narrower, more strongly falcate. Perichaetia with inner leaves gradually or suddenly narrowed to apex, plicate, margins denticulate or strongly so distally, apex acuminate, laminal cells smooth.

  • Provided by: [C].Flora of North America @ efloras.org
    • Source: [
    • 1
    • ]. 

     

    1. Cratoneuron filicinum (Hedw.) Spruce   牛角藓   niu jiao xian Cat. Musc. 21. 1867. Amblystegium filicinum (Hedw.) De Not., Comment. Soc. Crittog. Ital. 2: 291. 1867. Hypnum filicinum Hedw., Sp. Musc. Frond. 285. 1801. Type: Austria. Amblystegium campyliopsis Dixon, Hong Kong Naturalist, Suppl. 2: 24. f. 14. 1933. Type: China. Gansu province, Licent 250 (holotype BM). Amblystegium cuspidarioides (Müll. Hal.) Broth. in Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanocladus cuspidarioides (Müll. Hal.) Broth. ex Paris, Coll. Nom. Broth. 10: 1909. Drepanophyllaria cuspidarioides Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 5: 204. 1898. Type: China. Shaanxi (Schen-si), Liu-hua-zae, Giraldi 2092 (isotype BM). Amblystegium elegantifolium (Müll. Hal.) Broth. in Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanophyllaria elegantifolia Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 114. 1896. Type: China. Shaanxi, Aug. 1894, Giraldi s.n. Amblystegium fallax (Brid.) Milde, Bryol. Siles. 325. 1869. Cratoneuron filicinum var. fallax (Brid.) G. Roth, Eur. Laubm. 2: 532. 1904. Hypnum fallax Brid., Muscol. Recent. 2(2): 66. 1801. Amblystegium nivicalyx (Müll. Hal.) Broth. ex Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanophyllaria nivicalyx Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 115. 1896. Type: China. Shaanxi, Giraldi s.n. Amblystegium relaxum Cardot & Thér. in Thér., Bull. Acad. Int. Géogr. Bot. 3 sér., 15: 40. 1906. Type: China. Beijing (Pekin), May 1889, Bodinier s.n. (holotype PC). Amblystegium robustifolium (Müll. Hal.) Broth. ex Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanophyllaria robustifolia Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 5: 203. 1898. Type: China. Shaanxi (Schen-si), Giraldi 1877, 2096 (syntypes BM). Amblystegium tenax fo. spinifolium (Schimp.) Demaret, Bull. Jard. Bot. Etat 17: 363. 1945. Hygroamblystegium tenax var. spinifolium (Schimp.) Jenn., Man. Mosses W. Pennsylvania 278. 1913. Amblystegium irriguum var. spinifolium Schimp., Syn. Musc. Eur. (ed. 2) 713. 1876. Cratoneuron formosanum Broth., Ann. Bryol. 1: 22. 1928. Type: China. Taiwan, Taityu, Onoe, Suzuki 2894. Cratoneuron formosicum Broth. ex Sakurai, Bot. Mag. (Tokyo) 55: 210. 1941, hom. illeg. Cratoneuron longicostatum X.-L. Bai, Fl. Bryophyt. Intramongol. 385. 1997. Type: China. Nei Mongol, Mt. Helan, Z.-G. Tong 1249 (holotype HIMC; isotype IFSBH). Cratoneuron taihangense J.-C. Zhao, X.-Q. Li & L.-F. Han, J. Hebei Normal Univ. (Nat. Sci.) 25(1): 104. 2001. Type: China. Hebei, Pingshan Co., Qiandadi Mt., ca. 1450 m, 29 Feb. 2000, X.-Q. Li 20239 (isotype NY). Hygroamblystegium ramulosum Dixon in Yang, Sci. Rep. Natl. Tsing Hua Univ., Ser. B, Biol. Sci. 2: 129. 1936, nom. nud. Hypnum sinensimolluscum var. tenuius Müll. Hal. in Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 266. 1906, nom. nud. Thuidium leptopteris (Müll. Hal.) Paris, Index Bryol. 1284. 1898. Haplocladium leptopteris Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 116. 1896. Type: China. Shaanxi, Giraldi s.n.     1a. Cratoneuron filicinum var. filicinum   牛角藓原变种   niu jiao xian yuan bian zhong     Plants slender, medium-sized or large, green or yellowish green. Stems erect or erect-ascending, pinnately branched, sometimes irregularly pinnately branched, brownish rhizoids always present; paraphyllia foliose. Stem leaves erect or spreading, slightly curved, broadly ovate or ovate-lanceolate, 0.8–1.2 mm × 0.4–0.8 mm, abruptly acuminate above; margins usually serrulate; costae thick, ending in the leaf apex; median leaf cells oblong-rhomboidal, 20–35 µm × 4–7 µm, thin-walled; alar cells distinctly differentiated, strongly convex, extending close to costa at leaf base. Dioicous. Setae 30–40 mm long, reddish brown; capsules oblong-ellipsoidal to cylindrical, reddish brown. Spores 15–18 mm in diameter, finely papillose.  

  • Provided by: [B].Moss Flora of China
    • Source: [
    • 2
    • ]. 

    Plants medium-sized, rarely smaller; green, yellowish green, or occasionally brownish, often with pale shoot and branch apices. Stem somewhat stiff; pinnately or irregularly branched, occasionally almost unbranched; pseudoparaphyllia rounded to triangular; paraphyllia lanceolate or ovate-lanceolate, numerous or few, rarely absent in weak shoots; axillary hairs weak and infrequent, not more than one per axil, with 1-2-celled upper part, this hyaline; rhizoids frequently strongly branched, abundant, forming tomentum. Stem leaves straight or falcate (gradually curved), narrow- to broad-triangular or rounded-triangular to ovate, narrowing gradually to rather abruptly to acumen, not or hardly plicate, plane or slightly concave, decurrent; margin denticulate or serrulate almost throughout, rarely entire; costa strong, (42.0-)52.5-126.0 µm wide near base (outside the area to 157.0 µm), ending in leaf apex or excurrent, rarely ending slightly below leaf apex; median laminal cells 12.0-62.0 × 4.0-12.5 µm, somewhat incrassate, eporose; alar cells differentiated, usually numerous, strongly widened, hyaline; alar group well defined, transverse-triangular, widest toward leaf margin (except when modified by running water), reaching or nearly reaching costa. Branch leaves usually narrower and more falcate than stem leaves. Inner perichaetial leaves narrowing gradually or ± abruptly to acuminate apex. Exostome outside cross-striolate in lower part. [Sporophytes not known from neotropical material.]

  • Provided by: [F].Flora Neotropica
    • Source: [
    • 4
    • ]. 

    Plants medium-Sized to fairly robust, soft or rather rigid, extensive cushions up to about 12 cm high but usually considerably less, green, yellowish, or brownish, somewhat radiculose below and radiculose in tufts well up the stems. Stems spreading or, more often crowded and erect-ascending, irregularly or, more often, pinnately branched; paraphyllia usually none or very few but sometimes fairly numerous, variously lanceolate. Stem leaves erect, erect-spreading, or sometimes secund or falcate-secund, ± striate, 1-1.8 mm long, ovate-lanceolate to broadly ovate, rather abmptly acuminate; margins often somewhat recurved at the extreme base, serrulate nearly all around (in the acumen ±entire); costa rather strong, percurrent or ending in the acumen somewhat below the apex, rarely filling the acumen; cells smooth, firm-walled, shortly oblong-rhomboidal, about 3-5:1; alar cells abmptly differentiated and hyaline or yellow-brown, thin-or thick-walled in concave groups. Branch leaves shorter and narrower, usually ± falcate-secund. Setae 25-35 mm long; capsules 2-2.5 mm long; operculum convex-conic, apiculate or short-rostrate; cilia in 3s and 4s. Spores 15-18 µm, very finely roughened.

  • Provided by: [E].Memoirs of the New York Botanical Garden
    • Source: [
    • 9
    • ]. 

    Morphology

    Plants somewhat stiff. Stems with shoot and branch apices often pale; paraphyllia sometimes present only in youngest shoot portions; axillary hairs few, weak. Stem leaves gradually or suddenly narrowed to apex, plane or slightly concave; alar cells usually many, region large, usually strongly widened marginally, reaching from margin to costa or almost so; medial laminal cells 12-55 x 4-11 µm. Branch leaves usually narrower, more strongly falcate. Perichaetia with inner leaves gradually or suddenly narrowed to apex, plicate, margins denticulate or strongly so distally, apex acuminate, laminal cells smooth.

  • Provided by: [D].e-Flora of South Africa
    • Source: [
    • 3
    • ]. 

    Habitat

    Moist or wet habitats in calcareous districts, rock, tree bases beside streams, springs, moist soil, calcareous fens.

  • Provided by: [D].e-Flora of South Africa
    • Source: [
    • 3
    • ]. 

    Distribution

    Greenland; Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., Que., Sask., Yukon; Ala., Alaska, Ariz., Ark., Calif., Colo., Idaho, Ill., Ind., Iowa, Mich., Minn., Mo., Mont., Nev., N.H., N.Mex., N.Y., N.C., N.Dak., Ohio, Oreg., Pa., S.Dak., Tenn., Tex., Utah, Wash., Wis., Wyo.; Mexico (Baja California, México, Oaxaca, San Luis Potosí); South America; Eurasia; Africa; Atlantic Islands; Pacific Islands (New Zealand).

  • Provided by: [D].e-Flora of South Africa
    • Source: [
    • 3
    • ]. 

    In calcareous seepage; Baja California, Mexico, Oaxaca, San Luis Potosi.—Mexico; Guatemala; Newfoundland to Alaska, south to North Carolina, Texas and New Mexico; Europe, Asia, Japan; reported from Ecuador, North Africa and New Zealand.

  • Provided by: [E].Memoirs of the New York Botanical Garden
    • Source: [
    • 9
    • ]. 
    Flora NeotropicaGeneral Information

    Plants medium-sized, rarely smaller; green, yellowish green, or occasionally brownish, often with pale shoot and branch apices. Stem somewhat stiff; pinnately or irregularly branched, occasionally almost unbranched; pseudoparaphyllia rounded to triangular; paraphyllia lanceolate or ovate-lanceolate, numerous or few, rarely absent in weak shoots; axillary hairs weak and infrequent, not more than one per axil, with 1-2-celled upper part, this hyaline; rhizoids frequently strongly branched, abundant, forming tomentum. Stem leaves straight or falcate (gradually curved), narrow- to broad-triangular or rounded-triangular to ovate, narrowing gradually to rather abruptly to acumen, not or hardly plicate, plane or slightly concave, decurrent; margin denticulate or serrulate almost throughout, rarely entire; costa strong, (42.0-)52.5-126.0 µm wide near base (outside the area to 157.0 µm), ending in leaf apex or excurrent, rarely ending slightly below leaf apex; median laminal cells 12.0-62.0 × 4.0-12.5 µm, somewhat incrassate, eporose; alar cells differentiated, usually numerous, strongly widened, hyaline; alar group well defined, transverse-triangular, widest toward leaf margin (except when modified by running water), reaching or nearly reaching costa. Branch leaves usually narrower and more falcate than stem leaves. Inner perichaetial leaves narrowing gradually or ± abruptly to acuminate apex. Exostome outside cross-striolate in lower part. [Sporophytes not known from neotropical material.]

    Flora of North America @ efloras.orgGeneral Information

    Plants somewhat stiff. Stems with shoot and branch apices often pale; paraphyllia sometimes present only in youngest shoot portions; axillary hairs few, weak. Stem leaves gradually or suddenly narrowed to apex, plane or slightly concave; alar cells usually many, region large, usually strongly widened marginally, reaching from margin to costa or almost so; medial laminal cells 12-55 × 4-11 µm. Branch leaves usually narrower, more strongly falcate. Perichaetia with inner leaves gradually or suddenly narrowed to apex, plicate, margins denticulate or strongly so distally, apex acuminate, laminal cells smooth.

    Memoirs of the New York Botanical GardenDistribution

    In calcareous seepage; Baja California, Mexico, Oaxaca, San Luis Potosi.—Mexico; Guatemala; Newfoundland to Alaska, south to North Carolina, Texas and New Mexico; Europe, Asia, Japan; reported from Ecuador, North Africa and New Zealand.

    General Information

    Plants medium-Sized to fairly robust, soft or rather rigid, extensive cushions up to about 12 cm high but usually considerably less, green, yellowish, or brownish, somewhat radiculose below and radiculose in tufts well up the stems. Stems spreading or, more often crowded and erect-ascending, irregularly or, more often, pinnately branched; paraphyllia usually none or very few but sometimes fairly numerous, variously lanceolate. Stem leaves erect, erect-spreading, or sometimes secund or falcate-secund, ± striate, 1-1.8 mm long, ovate-lanceolate to broadly ovate, rather abmptly acuminate; margins often somewhat recurved at the extreme base, serrulate nearly all around (in the acumen ±entire); costa rather strong, percurrent or ending in the acumen somewhat below the apex, rarely filling the acumen; cells smooth, firm-walled, shortly oblong-rhomboidal, about 3-5:1; alar cells abmptly differentiated and hyaline or yellow-brown, thin-or thick-walled in concave groups. Branch leaves shorter and narrower, usually ± falcate-secund. Setae 25-35 mm long; capsules 2-2.5 mm long; operculum convex-conic, apiculate or short-rostrate; cilia in 3s and 4s. Spores 15-18 µm, very finely roughened.

    e-Flora of South AfricaMorphology

    Plants somewhat stiff. Stems with shoot and branch apices often pale; paraphyllia sometimes present only in youngest shoot portions; axillary hairs few, weak. Stem leaves gradually or suddenly narrowed to apex, plane or slightly concave; alar cells usually many, region large, usually strongly widened marginally, reaching from margin to costa or almost so; medial laminal cells 12-55 x 4-11 µm. Branch leaves usually narrower, more strongly falcate. Perichaetia with inner leaves gradually or suddenly narrowed to apex, plicate, margins denticulate or strongly so distally, apex acuminate, laminal cells smooth.

    Habitat

    Moist or wet habitats in calcareous districts, rock, tree bases beside streams, springs, moist soil, calcareous fens.

    Distribution

    Greenland; Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., Que., Sask., Yukon; Ala., Alaska, Ariz., Ark., Calif., Colo., Idaho, Ill., Ind., Iowa, Mich., Minn., Mo., Mont., Nev., N.H., N.Mex., N.Y., N.C., N.Dak., Ohio, Oreg., Pa., S.Dak., Tenn., Tex., Utah, Wash., Wis., Wyo.; Mexico (Baja California, México, Oaxaca, San Luis Potosí); South America; Eurasia; Africa; Atlantic Islands; Pacific Islands (New Zealand).

    Moss Flora of ChinaGeneral Information

     

    1. Cratoneuron filicinum (Hedw.) Spruce   牛角藓   niu jiao xian Cat. Musc. 21. 1867. Amblystegium filicinum (Hedw.) De Not., Comment. Soc. Crittog. Ital. 2: 291. 1867. Hypnum filicinum Hedw., Sp. Musc. Frond. 285. 1801. Type: Austria. Amblystegium campyliopsis Dixon, Hong Kong Naturalist, Suppl. 2: 24. f. 14. 1933. Type: China. Gansu province, Licent 250 (holotype BM). Amblystegium cuspidarioides (Müll. Hal.) Broth. in Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanocladus cuspidarioides (Müll. Hal.) Broth. ex Paris, Coll. Nom. Broth. 10: 1909. Drepanophyllaria cuspidarioides Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 5: 204. 1898. Type: China. Shaanxi (Schen-si), Liu-hua-zae, Giraldi 2092 (isotype BM). Amblystegium elegantifolium (Müll. Hal.) Broth. in Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanophyllaria elegantifolia Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 114. 1896. Type: China. Shaanxi, Aug. 1894, Giraldi s.n. Amblystegium fallax (Brid.) Milde, Bryol. Siles. 325. 1869. Cratoneuron filicinum var. fallax (Brid.) G. Roth, Eur. Laubm. 2: 532. 1904. Hypnum fallax Brid., Muscol. Recent. 2(2): 66. 1801. Amblystegium nivicalyx (Müll. Hal.) Broth. ex Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanophyllaria nivicalyx Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 115. 1896. Type: China. Shaanxi, Giraldi s.n. Amblystegium relaxum Cardot & Thér. in Thér., Bull. Acad. Int. Géogr. Bot. 3 sér., 15: 40. 1906. Type: China. Beijing (Pekin), May 1889, Bodinier s.n. (holotype PC). Amblystegium robustifolium (Müll. Hal.) Broth. ex Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 247. 1906. Drepanophyllaria robustifolia Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 5: 203. 1898. Type: China. Shaanxi (Schen-si), Giraldi 1877, 2096 (syntypes BM). Amblystegium tenax fo. spinifolium (Schimp.) Demaret, Bull. Jard. Bot. Etat 17: 363. 1945. Hygroamblystegium tenax var. spinifolium (Schimp.) Jenn., Man. Mosses W. Pennsylvania 278. 1913. Amblystegium irriguum var. spinifolium Schimp., Syn. Musc. Eur. (ed. 2) 713. 1876. Cratoneuron formosanum Broth., Ann. Bryol. 1: 22. 1928. Type: China. Taiwan, Taityu, Onoe, Suzuki 2894. Cratoneuron formosicum Broth. ex Sakurai, Bot. Mag. (Tokyo) 55: 210. 1941, hom. illeg. Cratoneuron longicostatum X.-L. Bai, Fl. Bryophyt. Intramongol. 385. 1997. Type: China. Nei Mongol, Mt. Helan, Z.-G. Tong 1249 (holotype HIMC; isotype IFSBH). Cratoneuron taihangense J.-C. Zhao, X.-Q. Li & L.-F. Han, J. Hebei Normal Univ. (Nat. Sci.) 25(1): 104. 2001. Type: China. Hebei, Pingshan Co., Qiandadi Mt., ca. 1450 m, 29 Feb. 2000, X.-Q. Li 20239 (isotype NY). Hygroamblystegium ramulosum Dixon in Yang, Sci. Rep. Natl. Tsing Hua Univ., Ser. B, Biol. Sci. 2: 129. 1936, nom. nud. Hypnum sinensimolluscum var. tenuius Müll. Hal. in Levier, Nuovo Giorn. Bot. Ital., n.s., 13: 266. 1906, nom. nud. Thuidium leptopteris (Müll. Hal.) Paris, Index Bryol. 1284. 1898. Haplocladium leptopteris Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 116. 1896. Type: China. Shaanxi, Giraldi s.n.     1a. Cratoneuron filicinum var. filicinum   牛角藓原变种   niu jiao xian yuan bian zhong     Plants slender, medium-sized or large, green or yellowish green. Stems erect or erect-ascending, pinnately branched, sometimes irregularly pinnately branched, brownish rhizoids always present; paraphyllia foliose. Stem leaves erect or spreading, slightly curved, broadly ovate or ovate-lanceolate, 0.8–1.2 mm × 0.4–0.8 mm, abruptly acuminate above; margins usually serrulate; costae thick, ending in the leaf apex; median leaf cells oblong-rhomboidal, 20–35 µm × 4–7 µm, thin-walled; alar cells distinctly differentiated, strongly convex, extending close to costa at leaf base. Dioicous. Setae 30–40 mm long, reddish brown; capsules oblong-ellipsoidal to cylindrical, reddish brown. Spores 15–18 mm in diameter, finely papillose.  

    Synonyms

     Information From

    Amblystegiaceae
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    World Flora Online Data. 2022.
    • A CC0 1.0 Universal (CC0 1.0).
    Moss Flora of China
    World Flora Online Data. 2024.
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    Flora of North America @ efloras.org
    World Flora Online Data. 2024.
    • C Flora of North America Association
    e-Flora of South Africa
    e-Flora of South Africa. v1.36. 2022. South African National Biodiversity Institute. http://ipt.sanbi.org.za/iptsanbi/resource?r=flora_descriptions&v=1.36
    • D All Rights Reserved
    Memoirs of the New York Botanical Garden
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    World Flora Online consortium
    http://www.worldfloraonline.org/organisation/WFO
    World Flora Online Data. 2024.
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