The World Flora Online

Herbs, perennial, to 45 cm; rhizomes absent; stolons present; corms present. Leaves emersed; petiole triangular, erect to ascending, 6.5--51 cm; blade sagittate, rarely hastate, 1.5--30.5 ´ 2--17 cm, basal lobes equal to or less than remainder of blade. Inflorescences racemes, rarely panicles, of 3--9 whorls, emersed, 4.5--28.5 ´ 4--23 cm; peduncles 10--59 cm; bracts connate more than or equal to ¼ total length, elliptic to lanceolate, 3--8 mm, delicate, not papillose; fruiting pedicels spreading, cylindric, 0.5--3.5 cm. Flowers to 4 cm diam.; sepals recurved to spreading, not enclosing flower or fruiting head; filaments cylindric, longer than anthers, glabrous; pistillate pedicellate, without ring of sterile stamens. Fruiting heads 1--1.7 cm diam; achenes oblanceoloid, without abaxial keel, 2.5--3.5 ´ to 2 mm, beaked; faces not tuberculate, wings absent, glands (0--)1(--2); beak lateral, horizontal, 1--2 mm. 2n = 22.

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4. Sagittaria latifolia Willd., Sp. Pl. 4: 409 (1805). Tipo: Canadá, Muhlenberg s.n. (US).

Por A. Lot H. y A. Novelo R.

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Sagittaria latifolia Willd., Sp. Pl. 4: 409. 1805; S. hastata Pursh; S. gracilis Pursh; S. esculenta Howell; S. obtusa Muhl. ex Willd.; S. ornithorhyncha Small; S. planipes Fernald; S. simplex Pursh; S. variabilis Engelm.; Urospatha friedrichsthalii Schott.

Hierbas perennes, glabras o pubescentes, hasta 45 cm de alto, con rizomas erectos y/o estolones; rizomas hasta 2 cm de largo y 0.5–1 cm de diámetro, estolones hasta 30 cm de largo y 0.3 cm de diámetro. Hojas emergentes, sagitadas o raramente hastadas, 10–30.5 cm de largo y 3.3–17 cm de ancho, ápice agudo o raramente obtuso, con 7–13 nervios, los lobos basales 4.5–16.5 cm de largo, separados entre sí 3.5–23 cm, con 4–6 nervios; pecíolos triangulares, 23.5–51 cm de largo, con una vaina 8–17.5 cm de largo en la base. Inflorescencia un escapo simple o raramente paniculado, con 3–9 verticilos, 13–28.5 cm de largo y 4–23 cm de ancho, los verticilos con 3 flores, pedúnculos triangulares, 32.5–59 cm de largo, brácteas lanceoladas, 4–8.4 mm de largo y 6.4–7 mm de ancho, delicadas a ligeramente gruesas, glabras, con el ápice obtuso a ligeramente agudo, pedicelos patentes a ascendentes cuando en flor y fruto, cilíndricos, 1.4–2.6 cm de largo y 1.1–1.4 mm de diámetro, flores unisexuales; sépalos reflexos a patentes, 5.5–9 mm de largo y 3–7 mm de ancho, glabros; pétalos unguiculados, 0.9–1.4 cm de largo y 0.8–1.9 cm de ancho, la uña 1–1.5 mm de largo; flores estaminadas con filamentos lineares, 1–3 mm de largo, glabros, anteras 1.2–1.9 mm de largo; flores pistiladas sin anillo de estambres estériles. Fruto rostrado, 2.5–3.5 mm de largo y hasta 2 mm de ancho, márgenes enteros o ligeramente dentados, caras sin alas y con (0–) 1 (–2) glándulas; rostro lateral, horizontal, 1–2 mm de largo.

Conocida en Nicaragua por una sola colección (Moreno 1628) de áreas inundadas y a lo largo de los márgenes de caños y lagos, Nueva Segovia; 500–700 m; fl y fr ago; ampliamente distribuida desde Canadá hasta Ecuador.

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Perennes, provenientes de rizomas o cormos,monoicas o dioicas. Hojas erectas, con pecíolo hastaca. 70 cm, la lámina 12–35 3–30 cm, linear a am-pliamente ovada, sagitada en la base. Infl. bisexual o (frecuentemente) unisexual, emergente, con 3–9 vertici-los simples (rara vez los inferiores ramificados), cada uno con 3 fls. Fls. con estambres 16–18 (fls.estaminadas), sin estambres estériles en las fls. pistiladas; pistilos estériles ausentes en las fls. estaminadas.Cabeza frutal 10–13 10–15 mm; pedicelo ascendente; sépalos reflexos; aquenios 2.5–3.5 mm, la cara lat-eral lisa o con 1(2) glándula, el pico lateral, 1–2 mm, horizontal.

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Plant glabrous or variously pubescent. Perennial with rhizomes and/or runners and corms. Leaves emersed or submerged, the emersed occasionally linear to ovate, mostly sagittate with linear to ovate-triangular blades 3-26 x 0.7-26 cm, the basal lobes 2-24 x 0.5-15 cm, linear to ovate-triangular, the submerged rare, modified into linear phyllodia to 60 cm long, the petioles to 100 cm long. Scape 15-80 cm long, with 2-7(-10) whorls of flowers, occasionally branching from the lowest whorl. Bracts 0.4-0.9(-1.5) cm long, boat-shaped, joined at the base or nearly free. Pedicels ascending, 0.5-6 cm long. Sepals reflexed, 0.5-1.1 cm long. Petals white, ca. twice as long as the sepals. Stamens 21-[infinity], the glabrous more or less linear filaments 1-3.7 mm long, the oblong anthers 1-2.3 mm long. Mature receptacle 1-2.5 cm in diameter; achenes obovate, 2.5-4 x 1.5-2.5 mm, the occasionally crested dorsal wing 0.4-0.8 mm wide, the ventral wing somewhat narrower, the faces with a resin duct and/or a low rib, or frequently unadorned, the gradually tapering, laterally inserted beak 0.6-1.6 mm long.

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Vegetatively very plastic, ± erect and with large, edible tubers; lvs sagittate or sometimes narrow and phyllodial, the blade 5–40 × 0.5–25 cm, with narrow or broad lobes; scape 1–12 dm, with 2–15 whorls of fls, sometimes branching at the lowest whorl, the upper fls staminate on short pedicels, the lower pistillate on longer, ± ascending pedicels; bracts 4–15 mm, boat-shaped, broad-based, rounded above or broadly acute, 4–15 mm, papery but friable at the tip; sep ovate, obtuse, 5–11 mm, reflexed in fr; pet 1–2 cm; stamens 20–40, on slender, glabrous filaments; achenes 2.5–4 mm, winged on the margins only, or with a poorly developed wing on each face, with (var. latifolia) or without (var. pubescens) resin-ducts; beak 0.6–1.8 mm, set at essentially right angles to the body; 2n=22. Abundant in swamps, ponds, and streams; N.S. and Que. to s. B.C., s. to trop. Amer. July–Sept. The widespread, glabrous var. latifolia is uncommon in the principal range of the stellate-hairy var. pubescens (Muhl.) J. G. Sm., which occurs chiefly from the mts. to the coast, from s. Pa. to W.Va. and Ga., w. occasionally to O., Tenn., and Tex.

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Perennial herbs from stolons and/or corms, glabrous or pubescent, to 45 cm tall; stolons to 30 cm long, to 0.3 cm wide; corms 0.5-1 cm long, to 2 cm diam. Leaves emersed petiolate, the petioles triangular, 6.5-51 cm long, 2-8 mm wide, the basal sheath to 1.5-17.5 cm, the blades pale green, sagittate or rarely hastate, 1.5-30.5 cm long, 2-17 cm wide, the veins 7-13, the apex acute or rarely obtuse, the basal lobes present, 0.5-16.5 cm long, the tips separated by 1-23 cm, with 2-6 veins. Inflorescence a simple or rarely paniculate scape of 3-9 whorls, emersed, 4.5-28.5 cm long, 4-23 cm wide, the whorls with 3 flowers; peduncles triangular, 10-59 cm long, 0.5-0.7 mm diam.; staminate bracts united, elliptic, delicate, 3-5.5 mm long, the apex obtuse; carpellate bracts separate, lanceolate, delicate, 5.5-8 mm long, the apex obtuse; staminate pedicels spreading, cylindric, 0.8-2.5 cm long, 0.1-0.4 mm diam.; carpellate pedicels spreading in flower and fruit, cylindric, 0.5-3.5 cm long, 0.5-1 mm diam. Staminate flowers without sterile carpels, the sepals reflexed to spreading, 5-6 mm long, 3-4.5 mm wide, the petals clawed, 6-9 mm long, 4.5-5 mm wide, the claws ca. 1.5 mm long, the stamens 16-18, the filaments glabrous, cylindric, 1-3 mm long, 0.2-0.3 mm wide, the anthers linear, 1.2-1.9 mm long, 0.5-0.6 mm wide, the apex obtuse; carpellate flowers without ring of sterile stamens, the sepals reflexed in flower and fruit, ca. 1 cm long, ca. 0.8 cm wide, the petals clawed, ca. 18 mm long, ca. 12 mm wide, the claw ca. 0.5 mm long. Fruit aggregate 1-1.7 cm diam.; achenes oblanceolate, beaked, without keel, 2.5-3.5 mm long, to 2 mm wide, the faces etuberculate, with (0-)l(-2) glands, the beak lateral, horizontal, 1-2 mm long.

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Ähnlich wie S. sagittifolia, aber kräftiger, Blätter mit 5–12 cm breiten Abschnitten, der mittlere oft stumpf, Blattstiel rundlich (bei S. sagittifolia scharf 3eckig), Blütendurchmesser 3–4 cm (bei S. sagittifolia 1,5–2,

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7–9. Gräben, Ufer. Aus Nordamerika stammende Aquarienpflanze, da und dort ausgepflanzt und verwildert, kollin. M.

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Throughout North America, south through Mexico, Central America, and northwestern South America.

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